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Gökdeniz and Rubin Kazan stun reigning champion Barca
Sevilla made it three wins in a row with a 3-1 success at VfB Stuttgart but Arsenal were denied their third successive win by a last-minute equaliser for AZ Alkmaar in a 1-1 draw.
Barcelona fell behind after two minutes at the Nou Camp when Aleksandr Ryazantsev scored from long range. Zlatan Ibrahimovic equalized in the 48th minute but former Turkey international midfieldfer Gökdeniz Karadeniz restored the Kazan's lead 20 minutes from time and, despite a series of late attacks, Barca could not prevent their first defeat in the competition in 10 games.
The results leave Group F, which had looked a two-horse race at the start, too close to call with Barcelona, Kiev and Rubin all on four points and Inter on three after three draws.
An injury-time goal by Cesar Delgado put Lyon in control of Group E and left Liverpool's hopes hanging by a thread. Five-time European champion Liverpool, already struggling without injured striker Fernando Torres, lost injured captain Steven Gerrard after 24 minutes but went ahead through Yossi Benayoun late in the first half.
Lyon substitute Maxime Gonalons headed a deserved equaliser after 72 minutes before Argentine forward Delgado finished off a sharp breakaway to hand Liverpool a fourth successive defeat in all competitions and leave them third on just three points.
Sevilla are also on nine points after two goals from defender Sebastien Squillaci and one for Jesus Nava gave them an impressive Group G success in Germany.
Romanian champion Unirea Urziceni is the unlikely second-placed side on four points. It secured a first European win by embarrassing Rangers 4-1 in Glasgow -- helped by two own goals and a missed penalty from the generous host.
Arsenal also looked on course for a third successive win until David Mendes da Silva grabbed a stoppage time equalizer to earn AZ Alkmaar a point.
Arsenal remains top of Group H on seven points with Olympiakos second on six after it also scored in added time to beat Standard Liege 2-1.
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